Error thrown when closest() calls matchesSelector method
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Error is as follows: expr.replace is not a function
This occurs due to the closest method calling calling jQuery.find.matchesSelector() with improperly typed arguments. The second parameter (selectors), should be passed as a string, but is an object.
At least I think that's whats going on :D.
This offending line is 4424 in uncompressed 1.4.4.